Harlestone is a village and civil parish in the West Northamptonshire district, in the ceremonial county of Northamptonshire, England. The parish had a recorded population of 445 in the 2011 census. From 1974 to 2021 it was in Daventry district. The village is divided into two smaller settlements: Upper Harlestone and Lower Harlestone, which are linked by road, footpaths and bridleways. Lower Harlestone lies along the A428, and Upper Harlestone is directly to the south-west.